You have been given some good tips thus far,and I am far from accomplished at guitar,but some things that help me are.
Try to say more with less notes.Pick a box of any 4 notes in a scale for example and try to ring out as much expression from just those 4 notes only(think Billy Gibbons from ZZ Top,a master of wringing 4 notes dry).Another is to remember that silence at the right spot is just as powerful as a played note,so don't just keep a running flurry of notes.You wouldn't talk that way,unless you were manic or all coked up,and has anyone ever had a good conversation with someone who was either? Think of it as speaking with your guitar and you don't usually speak in a run on sentence,so don't fill every space with notes,let it breathe a little and let the listener digest what you just "said" with your guitar.
Finally I would say that you take some of the scales that you have learned,forget about them and don't think that you have to stay right in that one scale pattern only.Don't be afraid to go out in any direction that may or may not be in that scale mode or not.Let your ears dictate what step you need to take,and even if it is wrong,play it with rythmic conviction and think about your overall solo in terms of rythm and what the song's rythm is and staying within it and noone but the most seasoned of ears will tell if your playing a "wrong"note or not.Plus there are really no "wrong" notes,just ones not played with conviction.
One more thing,alot of times when you have a rythm down cold,just close your eyes and let go and don't think about what you want to play or where you are going with it,just feel the music and the vibration and it will lead you.The better in tune with it you get,the better your solo's will be.But I know that is a sucky esoteric what the hell kind of metaphysical B.S. are you talking about answer,but believe me it will make sense.The more we do anything the better we get at it.
These are a few things that have helped me get to the point that I am,but like I wrote earlier,I still have a long ways to go too,and this is a neverending lifetime journey,so really try to enjoy the process.